Patrick McMillen received a B.S. in Biology from Tufts University and his Ph.D. from the Yale University Department of Molecular, Cellular and Developmental Biology.

Patrick McMillen studied Biology as an undergraduate at Tufts before pursuing his Ph.D. in the Yale Department of Molecular, Cellular and Developmental Biology and ultimately returning to Tufts to join the Levin Laboratory. He has worked extensively with zebrafish and currently works with frogs. At heart a developmental biologist, Patrick studies the ways that groups of cells collaborate to become greater than their individual parts. He leverages cutting edge microscopy to better reveal the invisible world of bioelectricity and other paradigms of physiological computation, and is enthusiastic about collaborating with other researchers to more clearly see these things in their own systems.